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Kingdom: AnimaliaPhylum: ChordataClass: MammaliaOrder: CarnivoraFamily: Canidae (dog family)Genus: Canis (Latin word meaning \"dog\")Species: lupus (Greek word meaning \"wolf\")Common names: gray wolf, timber wolfNames in other languages: Lobo (Spanish), Loup (French), Lupo (Italian), Varg (Swedish), Ulv (Norwegian)Physical CharacteristicsAverage body mass: males 110 pounds (50 kg); females 90 pounds (41 kg)Heaviest known wolf in Yellowstone: 148 pounds (wolf 760M of Yellowstone Delta pack with no food in stomach)Average height at shoulder: males 81 cm, females 77 cmAverage length: 181 cmEyes: blue at birth, light yellow to gold to brown as an adultNumber of bones: 319 males, 318 femalesNumber of teeth: 42Dental formulae: incisors 3 top/3 bottom, canines 1/1, premolars 4/4, molars 3/2 (on each side)Pelage: gray or black (ratio 50:50), rarely whiteBlack coat color: caused by K-locus gene thought to have originated from historic hybridization with domestic dogs 500-14,000 years agoLocomotion: tetrapedal, digitigradeAverage rate of speed: 5 miles/hour (8 kph)Top speed: 35 miles/hour (56 kph)Body temperature: 100-102.5 F (37.3-39.1 C)Respiration: 10-30 breathes per minuteHeart rate: 70-120 beats per minuteBite pressure: 1,200 psiSenses and CommunicationSmell: excellent, although unmeasured. Estimated to be thousands of times better than humansVision: excellent night vision; no red or green cones, but have blue and yellow conesHearing: little is known, but probably similar to dogs (relatively normal hearing abilities compared to other mammals)Howling function: many uses, including intrapack communication, advertising territory, coordinating social activitiesDistance howling can be heard: forest=11km (6.6 mi), open areas=16 km (9.6 mi)DietFeeding habits: generalist carnivore; scavenges when possible and has been known to eat small amounts of vegetationPrimary food sources in Yellowstone: Winter: elk (>96%), bison (3-4% and increasing in recent years; deer (1.5%); Spring: elk (89%), bison (7%), deer (7.1%); Summer: elk (85%), bison (14.1%), deer (5 years old: 18%Current North American population: 67,100-74,100 (53,600-57,600 of these in Canada)Average home range size in Yellowstone (northern range): 274 km2 (range=58-1,151 km2)Average home range size in Yellowstone (interior): 620 km2 (range=105-1675 km2)Average home range size in Yellowstone (park-wide): 428 km2Group of wolves: pack/ family (one of few eusocial species)Average pack size in Yellowstone: 9.8Largest pack recorded in Yellowstone: Druid Peak, 37 wolves (2001); may be the largest ever recorded (42 wolves seen together in Wood Buffalo National Park (1974) but unknown if they were a single pack)Percent of population that are lone wolves in Yellowstone: 2-5%Percent of population that are lone wolves in North America: 10-15%Sex ratio: 50:50Breeding and PupsMating: usually monogamous, but about 25% of packs have multiple breeding pairs under polygymous matingsCourtship: mid-FebruaryGestation: 63 daysBirth period: mid-AprilBirth location: denTypical dens: excavated under large roots, boulders, hillsides, caves with a tunnel leading to an enlarged chamber; several entrances and chambers may be presentDen emergence: 10-14 daysAverage litter size in Yellowstone: 4.4 at den emergence, 3.2 survive until late DecemberMaximum litter size recorded in Yellowstone: 11Split litters: multiple fathers per litter have not been detected in wild gray wolvesWeaning: 5-9 weeks from milk, then brought food (regurgitation) for another 3 monthsMilk content: 6.6% fat; 144 kCal per 100 gramsRendezvous sites: used as wolf pups get older as a central homesite; time spent there and number of homesites varies widely between packsAverage female age at first litter in Yellowstone: 2.7Oonset of female reproduction senescence: 4-5 yearsInterbirth interval: can be every yearEyes open: 12-14 daysDispersal: both sexes, YNP average age 2 years, 1 month; range 1-4 years
Yes, 2020 was a movie year like no other, and 2021 didn't exactly represent a glorious comeback. But despite the closure of cinemas, the switch to streaming and wild uncertainty over what comes next, there were plenty of mesmerising films to enjoy. Last year was a great one for fans of Spielberg, Jane Campion and more, while an iconic actor bowed out of an even more iconic role. For proof, just check out our pick of the top 20 films of 2021. Stock up on snacks, and let the movie marathons begin. 153554b96e
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